I mean sites like Shopify, Hulu, YellowPages.com, etc. "handle the strain" just fine.
Other components such as the search were probably also quite tricky. One thing i've never figured out is how Facebook handles search on your timeline. That a seriously complex problem.
Linkedin Recently published a bunch of papers about their feed tech:
https://engineering.linkedin.com/blog/2016/03/followfeed--li...
And Stream's stackshare is also interesting: https://stackshare.io/stream/stream-and-go-news-feeds-for-ov...
[1] https://www.wired.com/2015/09/whatsapp-serves-900-million-us...
From a user's point of view, they don't. I search for things a lot (or at least, I used to before I realized how bad it is) and things that I know I saw yesterday don't come up - even if I put in a very specific search. Sometimes it'll say "no results found", but I'll find it in a tab I hadn't closed yet.
They could handle the strain, within what their audience considered acceptable. That's the important thing. The key to scaling a startup is to always be just ahead of where your audience gives up. Anything less and your precious users will abandon ship and you'll fail; anything more you're burning too much cash, you'll run out of runway and you'll fail.