Cross-plane optical links would have a trickier tracking problem.
While there's no explicit mention of same-plane vs cross-plane optical links, I assume that the first time people have a public cross-plane optical link, they will make a big deal out of it. :)
The article also mentions that SpaceX would need to do further study before using laser links between satellites and ground stations-- this kind of optical link would require both more angular tracking and probably atmospheric correction as well.
And if they have zones where they don’t go to adjacent orbits, but instead go up or down within their orbit for the handover between orbits.
I believe Iridium had way more downlinks than they used to pre-bankruptcy. I guess volume constraints were less of an issue, so ok to hop around more in space.